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What are thistles?

What are thistles? Here are some definitions.

Noun
  1. plural of thistle
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In the winter, Chukars feed primarily on seeds, cheatgrass, and thistles, switching to insects and green leaves in the summer.
The yampas, glacier lilies, biscuit roots, and thistles are among her favorite flower roots.
But just let it rain and the next thing I know I've got thistles out there with stems big as my wrist.
For instance, dock and beggarticks often indicate wet soil, while thistles and mullein indicate a dry soil.
Two years ago there were lovely flowers growing here and now, there's nothing but thistles and weeds.
The flower heads are small and numerous compared to other thistles and the roots are extensive.

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