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What is a rune?

What is a rune? Here are some definitions.

Noun
  1. A letter, or character, belonging to the written language of various ancient Germanic peoples, especially the Scandinavians and the Anglo-Saxons.
  2. A Finnish or Scandinavian epic poem, or a division of one, especially a division of the Kalevala.
  3. A letter or mark used as mystical or magic symbol.
  4. A verse or song, especially one with mystical or mysterious overtones; a spell or an incantation.
  5. (obsolete) A roun (secret or mystery).
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Examples
Furthermore, several rune alphabets or futharks have been left behind to puzzle scholars.
Oddly enough, I memorized the Anglo-Saxon rune alphabet in high school with a friend of mine so we could pass notes in geometry class.
Nearly all the stones were cut in perfect spheres each with a rune that marked the element that they represented.
They hadn't been walking for very long when the mage stopped at a plain, metal door with a rune carved heavily in its surface.
His brother had spent nearly all day and night trying to carve a simple rune into the face of a small amethyst.
She paused, staring at the back, one finger gently tracing the black rune that appeared to have been burned into it.

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