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What is a rune?

What is a rune? Here are some definitions.

Noun
  1. A letter, or character, belonging to the written language of various ancient Germanic peoples, especially the Scandinavians and the Anglo-Saxons.
  2. A Finnish or Scandinavian epic poem, or a division of one, especially a division of the Kalevala.
  3. A letter or mark used as mystical or magic symbol.
  4. A verse or song, especially one with mystical or mysterious overtones; a spell or an incantation.
  5. (obsolete) A roun (secret or mystery).
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If you compare the futhorc given above, you will see that Tolkien has altered some the rune shapes.
She paused, staring at the back, one finger gently tracing the black rune that appeared to have been burned into it.
One was using recordings I made of rune meditations and mixing bits from each rune into a word spell.
There was a Danish king of that name who was famed for his generosity, to whom the rune poem probably refers.
If I was as drunk as these bottles say I was, I couldn't have cast a rune, let alone a spell.
Every stone of the edifice was marked with a rune, coursing with magical strength.

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