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What is a rune?

What is a rune? Here are some definitions.

Noun
  1. A letter, or character, belonging to the written language of various ancient Germanic peoples, especially the Scandinavians and the Anglo-Saxons.
  2. A Finnish or Scandinavian epic poem, or a division of one, especially a division of the Kalevala.
  3. A letter or mark used as mystical or magic symbol.
  4. A verse or song, especially one with mystical or mysterious overtones; a spell or an incantation.
  5. (obsolete) A roun (secret or mystery).
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Examples
There was a Danish king of that name who was famed for his generosity, to whom the rune poem probably refers.
If I was as drunk as these bottles say I was, I couldn't have cast a rune, let alone a spell.
She paused, staring at the back, one finger gently tracing the black rune that appeared to have been burned into it.
Furthermore, several rune alphabets or futharks have been left behind to puzzle scholars.
Nearly all the stones were cut in perfect spheres each with a rune that marked the element that they represented.
One may also bury a piece of wood inscribed with a protection rune or script.

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