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What is a rune?

What is a rune? Here are some definitions.

Noun
  1. A letter, or character, belonging to the written language of various ancient Germanic peoples, especially the Scandinavians and the Anglo-Saxons.
  2. A Finnish or Scandinavian epic poem, or a division of one, especially a division of the Kalevala.
  3. A letter or mark used as mystical or magic symbol.
  4. A verse or song, especially one with mystical or mysterious overtones; a spell or an incantation.
  5. (obsolete) A roun (secret or mystery).
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Examples
One may also bury a piece of wood inscribed with a protection rune or script.
They hadn't been walking for very long when the mage stopped at a plain, metal door with a rune carved heavily in its surface.
There was a Danish king of that name who was famed for his generosity, to whom the rune poem probably refers.
Oddly enough, I memorized the Anglo-Saxon rune alphabet in high school with a friend of mine so we could pass notes in geometry class.
His brother had spent nearly all day and night trying to carve a simple rune into the face of a small amethyst.
If I was as drunk as these bottles say I was, I couldn't have cast a rune, let alone a spell.

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