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What is a rune?

What is a rune? Here are some definitions.

Noun
  1. A letter, or character, belonging to the written language of various ancient Germanic peoples, especially the Scandinavians and the Anglo-Saxons.
  2. A Finnish or Scandinavian epic poem, or a division of one, especially a division of the Kalevala.
  3. A letter or mark used as mystical or magic symbol.
  4. A verse or song, especially one with mystical or mysterious overtones; a spell or an incantation.
  5. (obsolete) A roun (secret or mystery).
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Examples
One was using recordings I made of rune meditations and mixing bits from each rune into a word spell.
Every stone of the edifice was marked with a rune, coursing with magical strength.
She paused, staring at the back, one finger gently tracing the black rune that appeared to have been burned into it.
They hadn't been walking for very long when the mage stopped at a plain, metal door with a rune carved heavily in its surface.
Furthermore, several rune alphabets or futharks have been left behind to puzzle scholars.
One may also bury a piece of wood inscribed with a protection rune or script.

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