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What is a rune?

What is a rune? Here are some definitions.

Noun
  1. A letter, or character, belonging to the written language of various ancient Germanic peoples, especially the Scandinavians and the Anglo-Saxons.
  2. A Finnish or Scandinavian epic poem, or a division of one, especially a division of the Kalevala.
  3. A letter or mark used as mystical or magic symbol.
  4. A verse or song, especially one with mystical or mysterious overtones; a spell or an incantation.
  5. (obsolete) A roun (secret or mystery).
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Examples
His brother had spent nearly all day and night trying to carve a simple rune into the face of a small amethyst.
If I was as drunk as these bottles say I was, I couldn't have cast a rune, let alone a spell.
Nearly all the stones were cut in perfect spheres each with a rune that marked the element that they represented.
Oddly enough, I memorized the Anglo-Saxon rune alphabet in high school with a friend of mine so we could pass notes in geometry class.
If you compare the futhorc given above, you will see that Tolkien has altered some the rune shapes.
One may also bury a piece of wood inscribed with a protection rune or script.

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