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What is a rachis?

What is a rachis? Here are some definitions.

Noun
  1. (botany) The main shaft of either a compound leaf, head of grain, or fern frond.
  2. (anatomy) The spine or the vertebrae of the spine.
  3. (ornithology) The central shaft of a feather.
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Examples
The basal leaves can be more than 10 cm long and have three to eleven leaflets along their rachis.
The bristle and eyelash consist of a rachis void of barbs, except at the innermost base.
Feathers, however bizarre or morphologically complex, consist essentially of a rachis, barbs, and barbules.
Affected leaflets abscise from a compound leaf, while leaflets without symptoms remain on the rachis.
Inflorescences are the terminal toothbrush type, with five to 70 pairs of flowers on a rachis approx. 35-50 mm long.
Filoplumes have a rachis, with barbs and barbules somewhere along them, and they arise from follicles.

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