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What are quills?

What are quills? Here are some definitions.

Noun
  1. plural of quill
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Examples
In one arm, he cradled a stack of books, and in the other, a leather satchel full of feather quills and dozens of tiny glass bottles.
For example, the porcupine is defenseless except for its quills, the deer vulnerable except for its speed.
The clothing is made from white caribou hides and sewn with sinew, using split bird quills on the seams.
By the way, they're born with soft quills, and the quills harden up within about six to eight hours after birth.
And there it was, in plain sight by the door, with a small knife, the kind used to sharpen quills, hanging next to it.
When the quills begin to loosen, the bird removes them and is then ready to care for the new feathers.

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