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What are quills?

What are quills? Here are some definitions.

Noun
  1. plural of quill
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Throughout our period various breeds of birds were used to supply different quills, including duck, goose, swan and pheasant.
For example, the porcupine is defenseless except for its quills, the deer vulnerable except for its speed.
We were especially careful in excluding specimens in molt by checking all specimens for the presence of feather quills.
The quills are so lightly fixed to the porcupine's body that they are easily detached and left imbedded in the attacker.
And there it was, in plain sight by the door, with a small knife, the kind used to sharpen quills, hanging next to it.
When the quills begin to loosen, the bird removes them and is then ready to care for the new feathers.

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