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What are quadrats?

What are quadrats? Here are some definitions.

Noun
  1. plural of quadrat
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Gods, the quadrats used by printers in throwing on the imposing stone, similar to the movement in casting dice.
A total of 199 quadrats were sampled in closed canopy areas.
In September and October 2012, we sampled sixty-three 200-m2 quadrats across the 6 study sites.
Forest quadrats showed a less pronounced seasonal decline in light availability.
During June, July, and August of 2010, divers used quadrats to estimate densities of crabs and worm tubes.
Territory quadrats were the vegetation quadrats sampled at the center of each of 88 territories.

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