What does pupping mean?

Looking for the meaning or definition of the word pupping? Here's what it means.

Verb
  1. present participle of pup
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As no mother-pup pairs were observed, these ledges are molting but not pupping ledges.
Breeding and pupping sea otters generally occur from Point of Arches to the south, with a large concentration of sea otters near Cape Alava.
Loss of southern pupping areas due to inadequate or highly variable ice conditions may reduce these species as polar bear prey.
This, it is hoped, will succeed in revealing migratory patterns, aggregating sites and pupping grounds.
These results are in agreement with a report from the Oregon coast of a location that is also a harbor seal molting but not pupping area.
Massachusetts is the southernmost point of known pupping areas along the Atlantic Coast.

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