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What are foundresses?

What are foundresses? Here are some definitions.

Noun
  1. plural of foundress
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Examples
She and her colleagues tracked the fortunes of 1,113 foundresses in 228 nests in southern Spain.
Are lone paper wasp foundresses mainly the result of sister mortality?
Gene expression in workers was more similar to foundresses, which show maternal care, than to queens and gynes, which do not.
Because P simillimus foundresses are closely related, subordinate females would still gain in indirect fitness.
Every little Italian convent has managed to get their foundresses raised to sainthood, haven't they?
Because each colony is founded by a single queen, within-colony relatedness is higher than in species with multiple foundresses.

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