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What are ctenidia?

What are ctenidia? Here are some definitions.

Noun
  1. plural of ctenidium
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Examples
Meanwhile, Pelseneer had come up with a classification of bivalves based on the structure of the ctenidia.
No ctenidia but pallial branchiae in a circle between mantle and foot.
A single pair of ctenidia is present instead of the two pairs in Nautilus.
No ctenidia are present, and gas exchange is through the mantle surface.
Two pairs of ctenidia, and two pairs of renal tubes without reno-pericardial apertures.
All cephalopods have one pair of unciliated ctenidia within the mantle cavity, with the exception of Nautilus, which has two pairs of ctenidia.

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