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What are callouses?

What are callouses? Here are some definitions.

Noun
  1. plural of callous
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Examples
His hands were thin, with long fingers unmarked by callouses, and his skin was too white to have been too often outside.
As the callouses on the palms gave evidence of recent hard work, he was set free along with me.
On my word the good wife and mother hasn't the kinks out of her fingers yet, nor the callouses from her hands, by Jove!
But there was a sparkle in his eyes, an ease of movement and callouses on his hands.
His hair was greasy and stringy, and his skin had developed brown callouses.
They were large and shapely, but the only callouses they could show were accusingly recent.

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