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What is a groundling?

What is a groundling? Here are some definitions.

Noun
  1. Any of various plants or animals living on or near the ground, as a benthic fish or bottom feeder, especially:
    1. The spined loach (Cobitis taenia), weather loach ( Misgurnus fossilis), or other member of the loaches.
    2. The ringed plover, Charadrius hiaticula.
  2. An audience member in the cheap section (usually standing; originally in Elizabethan theater).
  3. By extension, an individual of uncultivated or uncultured taste.
  4. One who is confined to the ground, especially:
    1. (military, slang) A soldier who fights on the ground or serves as ground crew, as opposed to a seaman, pilot, etc.
    2. (fantasy) A member of a race that lives primarily underground, such as a dwarf.
  5. (Abrahamic religions) Adam, before eating the apple of knowledge of good and evil (emphasizing his creation from the ground).
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Examples
This adaptability has proven unfortunate for many natives species introduced to vine cats, such as the scrapbeak groundling, driven to extinction within a mere three months of contact.
The two of them were naked, the groundling and his woman, they were not ashamed.
You cared about Oedipus and Hamlet because they were noble and you were a groundling.
He then enlisted in the air force and served as a groundling in North Africa, Sicily, and England.
Pumpkins are among the most imposing of all groundling growths.
Still, the folio Ben looks to publish will be well beyond the purse of most scholars, let alone a groundling.

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