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What does holdfast mean?

Looking for the meaning or definition of the word holdfast? Here's what it means.

Noun
  1. Something to or by which an object can be securely fastened.
  2. (zoology) A root-like structure that anchors aquatic sessile organisms, such as seaweed, other sessile algae, stalked crinoids, benthic cnidarians, and sponges, to the substrate.
  3. (medicine) Actinomycosis.
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What do you think has become of Captain Hill and his companions, Mr. holdfast?
Figure 2: Diagrams of Chondrus crispus showing its discoid holdfast and Furcellaria lumbricalis showing its rhizoidal holdfast.
Many aquatic bacteria produce an acidic mucopolysaccharide holdfast, which allows them to adhere tightly to rocks or other surfaces.
Samples were taken from the harvest for holdfast analysis from each of the landing sites.
At the base of the stalk is the holdfast, which cements the alga to the rock and prevents it from being washed away.
At the end of the holdfast is a tiny disc, which secretes an extremely powerful glue.

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